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SAT 语法题There is a special relationship between a conductor a

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SAT 语法题
There is a special relationship between a conductor and the members of an orchestra,when at its best can be creative and at its worst,destabilizing.
答案是把when 改成which,为什么?
In Alaska,the ground is riddled with ice wedge,many quite ancient,caused( by the cold earth cracks) and the cracks fill with water.
答案是把by改成when,不理解,
第一句which指代relationship.
第二句如果用by因该接sth.应该是caused by sth.所以是caused by the cold earth's cracking.所以是when,接个句子.还不明白可在线问我.